#goldbeter2



#reactions
v_1:
 $pool = {1.0}Cvar
  cell * vi
v_2:
 {1.0}Cvar = $pool
  cell *kd*Cvar
v_3:
 {1.0}Cvar = $pool
  cell * vd*Cvar*X / (Kd + Cvar)
v_4:
 $pool = {1.0}Mvar
  cell *V1* (1 - Mvar) / (1 + K1 - Mvar)
v_5:
 {1.0}Mvar = $pool
  cell * V2*Mvar / (K2 + Mvar)
v_6:
 $pool = {1.0}X
  cell *V3* (1 - X) / (1 + K3 - X)
v_7:
 {1.0}X = $pool
  cell * V4*X / (K4 + X)


#parameters
K1 = 0.005
K2 = 0.005
K3 = 0.005
K4 = 0.005
Kc = 0.5
Kd = 0.02
V2 = 1.5
V4 = 0.5
VM1 = 3.0
VM3 = 1.0
cell = 1.0
kd = 0.01
vd = 0.25
vi = 0.025
default_compartment = 1.0


#initial values
Cvar = 0.01
Mvar = 0.01
X = 0.01


#assignment rules
!F V1 = (VM1*Cvar)/(Kc + Cvar)
!F V3 = VM3*Mvar