begin name
gardner1
end name

begin reactions
reaction1 {1.0}$EXTERNAL = {1.0}C
reaction10 {1.0}Z = {1.0}C
reaction11 {1.0}Z = {1.0}Y
reaction12 {1.0}$EXTERNAL = {1.0}Y
reaction13 {1.0}Y = {1.0}$EXTERNAL
reaction2 {1.0}C = {1.0}$EXTERNAL
reaction3 {1.0}C = {1.0}$EXTERNAL
reaction4 {1.0}$EXTERNAL = {1.0}M
reaction5 {1.0}M = {1.0}$EXTERNAL
reaction6 {1.0}$EXTERNAL = {1.0}X
reaction7 {1.0}X = {1.0}$EXTERNAL
reaction8 {1.0}C + {1.0}Y = {1.0}Z
reaction9 {1.0}Z = {1.0}C + {1.0}Y
end reactions

begin rate equations
reaction1 = reaction1vi
reaction10 = reaction10alpha*reaction10d1*Z[t]
reaction11 = reaction11alpha*reaction11kd*Z[t]
reaction12 = reaction12vs
reaction13 = reaction13d1*Y[t]
reaction2 = (reaction2k1*C[t]*X[t])/(reaction2K5 + C[t])
reaction3 = reaction3kd*C[t]
reaction4 = (V1*(1 - M[t]))/(1 + reaction4K1 - M[t])
reaction5 = (reaction5V2*M[t])/(reaction5K2 + M[t])
reaction6 = (V3*(1 - X[t]))/(1 + reaction6K3 - X[t])
reaction7 = (reaction7V4*X[t])/(reaction7K4 + X[t])
reaction8 = reaction8a1*C[t]*Y[t]
reaction9 = reaction9a2*Z[t]
end rate equations

begin parameters
K6 = 0.3
V1p = 0.75
V3p = 0.3
reaction1vi = 0.1
reaction2k1 = 0.5
reaction2K5 = 0.02
reaction3kd = 0.02
reaction4K1 = 0.1
reaction5V2 = 0.25
reaction5K2 = 0.1
reaction6K3 = 0.2
reaction7K4 = 0.1
reaction7V4 = 0.1
reaction8a1 = 0.05
reaction9a2 = 0.05
reaction10alpha = 0.1
reaction10d1 = 0.05
reaction11kd = 0.02
reaction11alpha = 0.1
reaction12vs = 0.2
reaction13d1 = 0.05
Cell = 1.0
end parameters

begin initial conditions
C[0] = Ci
M[0] = Mi
X[0] = Xi
Y[0] = Yi
Z[0] = Zi
end initial conditions

begin initial values
Ci = 0.0
Mi = 0.0
Xi = 0.0
Yi = 0.0
Zi = 0.0
end initial values

begin assignment rules
V3 := V3p*M[t]
V1 := (V1p*C[t])/(K6 + C[t])
end assignment rules

begin function definitions
end function definitions

begin events
end events

begin process annotations
end process annotations

begin species annotations
C http://identifiers.org/interpro/IPR006670
end species annotations

begin units
end units