begin name
decroly1
end name

begin reactions
r1 {1.0}$EXTERNAL = {1.0}alpha
r2 {1.0}alpha = {50.0}beta
r3 {1.0}beta = {0.02}gamma
r4 {1.0}gamma = {1.0}$EXTERNAL
end reactions

begin rate equations
r1 = r1vKm1
r2 = (r2sigma1*alpha[t]*(1 + alpha[t])*(1 + beta[t])^2)/(r2L1 + (1 + alpha[t])^2*(1 + beta[t])^2)
r3 = (r3sigma2*beta[t]*(1 + r3d*beta[t])*(1 + gamma[t])^2)/(r3L2 + (1 + r3d*beta[t])^2*(1 + gamma[t])^2)
r4 = r4ks*gamma[t]
end rate equations

begin parameters
r1vKm1 = 0.45
r2L1 = 500000000.0
r2sigma1 = 10.0
r3L2 = 100.0
r3d = 0.0
r3sigma2 = 10.0
r4ks = 1.99
cell = 1.0
end parameters

begin initial conditions
alpha[0] = alphai
beta[0] = betai
gamma[0] = gammai
end initial conditions

begin initial values
alphai = 29.19988
betai = 188.8
gammai = 0.3367
end initial values

begin assignment rules
end assignment rules

begin function definitions
end function definitions

begin events
end events

begin process annotations
end process annotations

begin species annotations
end species annotations

begin units
end units