begin name
arnold6
end name

begin reactions
GAP2Ru5P {1.0}GAP = {0.6}Ru5P
GAP2sink {1.0}GAP = {1.0}$EXTERNAL
GAPDH {1.0}DPGA = {1.0}GAP
PGA2sink {1.0}PGA = {1.0}$EXTERNAL
PGAK {1.0}PGA + {1.0}$ATP = {1.0}$ADP + {1.0}DPGA
Ru5PK {1.0}Ru5P + {1.0}$ATP = {1.0}RuBP + {1.0}$ADP
RuBisCO {1.0}RuBP = {2.0}PGA
end reactions

begin rate equations
GAP2Ru5P = chloroplast*function1[GAP[t], GAP2Ru5PKm, GAP2Ru5PV]
GAP2sink = chloroplast*function1[GAP[t], GAP2sinkKm, GAP2sinkV]
GAPDH = chloroplast*function1[DPGA[t], GAPDHKm, GAPDHV]
PGA2sink = chloroplast*function1[PGA[t], PGA2sinkKm, PGA2sinkV]
PGAK = chloroplast*function2[PGAKVm, PGA[t], ATP, PGAKK1, PGAKK2]
Ru5PK = chloroplast*function2[Ru5PKVm, Ru5P[t], ATP, Ru5PKK1, Ru5PKK2]
RuBisCO = chloroplast*function1[RuBP[t], RuBisCOKm, RuBisCOV]
end rate equations

begin parameters
ADP = 0.82
ATP = 0.68
RuBisCOKm = 1.0
RuBisCOV = 3.78
PGAKVm = 11.75
PGAKK1 = 0.24
PGAKK2 = 0.39
GAPDHKm = 0.5
GAPDHV = 5.04
GAP2Ru5PKm = 0.84
GAP2Ru5PV = 3.05
Ru5PKVm = 8.0
Ru5PKK1 = 0.15
Ru5PKK2 = 0.059
PGA2sinkKm = 0.75
PGA2sinkV = 3.0
GAP2sinkKm = 5.0
GAP2sinkV = 0.1
chloroplast = 1.0
end parameters

begin initial conditions
DPGA[0] = DPGAi
GAP[0] = GAPi
PGA[0] = PGAi
Ru5P[0] = Ru5Pi
RuBP[0] = RuBPi
end initial conditions

begin initial values
DPGAi = 0.0011
GAPi = 0.02
PGAi = 2.4
Ru5Pi = 0.0501
RuBPi = 2.0
end initial values

begin assignment rules
Pi := 15 - ADP - GAP[t] - PGA[t] - Ru5P[t] - 2*(ATP + DPGA[t] + RuBP[t])
end assignment rules

begin function definitions
function2[Vm_,s1_,s2_,K1_,K2_] := (s1*s2*Vm)/((K1 + s1)*(K2 + s2))
function1[substrate_,Km_,V_] := (substrate*V)/(Km + substrate)
end function definitions

begin events
end events

begin process annotations
end process annotations

begin species annotations
end species annotations

begin units
end units