$\mathrm{F1}(\frac{\mathrm{vm11}\mathrm{prot}(\mathrm{asp}\mathrm{atp}-\frac{\mathrm{aspp}\mathrm{adp}}{\mathrm{keqak}})}{(\frac{\mathrm{k11}(1+\left(\frac{\mathrm{thr}}{\mathrm{k1thr}}\right)^{\mathrm{nak1}})}{1+\left(\frac{\mathrm{thr}}{\mathrm{alpha}\mathrm{k1thr}}\right)^{\mathrm{nak1}}}+\frac{\mathrm{k11}\mathrm{aspp}}{\mathrm{k1aspp}}+\mathrm{asp})(\mathrm{k1atp}(1+\frac{\mathrm{adp}}{\mathrm{k1adp}})+\mathrm{atp})}+\frac{\mathrm{vm13}\mathrm{prot}(\mathrm{asp}\mathrm{atp}-\frac{\mathrm{aspp}\mathrm{adp}}{\mathrm{keqak}})}{(1+\left(\frac{\mathrm{lys}}{\mathrm{k1lys}}\right)^{\mathrm{nak3}})(\mathrm{k13}(1+\frac{\mathrm{aspp}}{\mathrm{k13aspp}})+\mathrm{asp})(\mathrm{k13atp}(1+\frac{\mathrm{adp}}{\mathrm{k13adp}})+\mathrm{atp})})$ $\frac{\mathrm{vm2f}\mathrm{prot}(\mathrm{aspp}\mathrm{nadph}-\frac{\mathrm{asa}\mathrm{nadp}\mathrm{pi}}{\mathrm{k2eq}})}{(\mathrm{k2aspp}(1+\frac{\mathrm{asa}}{\mathrm{k2asa}})(1+\frac{\mathrm{pi}}{\mathrm{k2p}})+\mathrm{aspp})(\mathrm{k2nadph}(1+\frac{\mathrm{nadp}}{\mathrm{k2nadp}})+\mathrm{nadph})}$ $\frac{\mathrm{vm3f}\mathrm{prot}(\mathrm{asa}\mathrm{nadph}-\frac{\mathrm{hs}\mathrm{nadp}}{\mathrm{k3eq}})}{\frac{(1+\frac{\mathrm{asp}}{\mathrm{k3asp}})(1+\left(\frac{\mathrm{thr}}{\mathrm{k3thr}}\right)^{\mathrm{nhdh1}})}{1+\left(\frac{\mathrm{thr}}{\mathrm{alpha3}\mathrm{k3thr}}\right)^{\mathrm{nhdh1}}}(\mathrm{k3asa}+\mathrm{asa}+\frac{\mathrm{hs}\mathrm{k3asa}}{\mathrm{k3hs}})(\mathrm{k3nadph}(1+\frac{\mathrm{nadp}}{\mathrm{k3nadp}})+\mathrm{nadph})}$ $\frac{\mathrm{vm4f}\mathrm{prot}\mathrm{hs}\mathrm{atp}}{(1+\frac{\mathrm{lys}}{\mathrm{k4lys}})(\mathrm{atp}+\mathrm{k4atp}(1+\frac{\mathrm{hs}}{\mathrm{k4ihs}}))(\mathrm{hs}+\mathrm{k4hs}(1+\frac{\mathrm{thr}}{\mathrm{k4thr}})(1+\frac{\mathrm{atp}}{\mathrm{k4iatp}}))}$ $\frac{\mathrm{vm5}\mathrm{prot}\mathrm{hsp}}{\mathrm{hsp}+\mathrm{k5hsp}}$