$\frac{\mathrm{k3}}{\mathrm{R5}+\mathrm{c2}}$ $\frac{\mathrm{k2}+\mathrm{r4}\mathrm{c2}}{\mathrm{R3}+\mathrm{c2}}$ $\frac{\mathrm{k1}}{\mathrm{R3}+\mathrm{c2}}$ $\frac{\mathrm{r2}\mathrm{c2}}{\mathrm{R1}+\mathrm{c2}}$ $\frac{\mathrm{k3}}{\mathrm{R5}+\mathrm{c1}}$ $\frac{\mathrm{k2}+\mathrm{r4}\mathrm{c1}}{\mathrm{R3}+\mathrm{c1}}$ $\frac{\mathrm{k1}}{\mathrm{R3}+\mathrm{c1}}$ $\frac{\mathrm{r2}\mathrm{c1}}{\mathrm{R1}+\mathrm{c1}}$ $\mathrm{compartment}\mathrm{Phi3\_c1}(1-\mathrm{h1})$ $\mathrm{compartment}\mathrm{Phi3\_c2}(1-\mathrm{h2})$ $\mathrm{compartment}\mathrm{Jleak}$ $\mathrm{compartment}\mathrm{Jleak}$ $\mathrm{compartment}\frac{\mathrm{Vp}\mathrm{c1}^2}{\mathrm{Kp}^2+\mathrm{c1}^2}$ $\frac{\mathrm{compartment}\mathrm{Vp}\mathrm{c2}^2}{\mathrm{Kp}^2+\mathrm{c2}^2}$ $\mathrm{compartment}\mathrm{kf}\left(\frac{p\mathrm{h1}\mathrm{Phi1\_c1}}{\mathrm{Phi1\_c1}p+\mathrm{Phi\_minus1\_c1}}\right)^4$ $\mathrm{compartment}\mathrm{kf}\left(\frac{p\mathrm{h2}\mathrm{Phi1\_c2}}{\mathrm{Phi1\_c2}p+\mathrm{Phi\_minus1\_c2}}\right)^4$ $\frac{\mathrm{compartment}\mathrm{Phi1\_c1}\mathrm{Phi2\_c1}\mathrm{h1}p}{\mathrm{Phi1\_c1}p+\mathrm{Phi\_minus1\_c1}}$ $\frac{\mathrm{compartment}\mathrm{Phi1\_c2}\mathrm{Phi2\_c2}\mathrm{h2}p}{\mathrm{Phi1\_c2}p+\mathrm{Phi\_minus1\_c2}}$ $\mathrm{compartment}D(\mathrm{c2}-\mathrm{c1})$